Los Rastros Formation
Adapted from Wikipedia · Discoverer experience
The Los Rastros Formation is an important rock layer from the middle of the Carnian age found in northwestern Argentina. It is part of the Agua de la Peña Group and is located in the Ischigualasto-Villa Unión Basin. This formation is special because many fossils have been discovered there.
Scientists have found tracks made by ancient animals called archosaurs, along with fossils of many insects, fish, bivalves, crustaceans, and plants. These fossils help us learn about the world millions of years ago and what kinds of life existed at that time.
Geology
The Los Rastros Formation lies above the Chañares Formation and/or the Ischicuca Formation, and is covered by the Ischigualasto Formation. It is part of the Agua de la Peña Group in the Ischigualasto-Villa Unión Basin, and can be seen in the Ischigualasto Provincial Park, a World Heritage Site in Argentina. This formation, which can be up to 1,000 metres (3,300 ft) thick, is made of claystones, mudstones, and sandstones. It formed in a deltaic to lacustrine environment during a time when the land was changing after a big split in the earth’s surface.
In 2020, scientists found that the lower part of the Los Rastros Formation is about 234.47 ± 0.44 million years old. This shows that most of the layers above it were put down during or after the Carnian Pluvial Event.
Paleobiota
The Los Rastros Formation is a place where scientists have found many old fossils. The only body fossil of a four-legged animal found here is a poorly preserved amphibian called a temnospondyl.
Ichnofossils
This formation is famous for its animal footprints, especially near a place called Quebrada de Ischichuca. These footprints show animals that walked on two legs with three toes on the ground. Scientists think these might be early dinosaurs or similar animals. Some footprints also show hand prints, suggesting animals that walked on four legs. Other footprints might have been made by groups of small reptiles or even large four-legged animals moving together.
Fish
Invertebrates
Many insect fossils have been found here, including types like cockroaches, beetles, and true bugs. Other small animals found include clam shrimp and freshwater mussels.
Blattoptera
Coleoptera
Hemiptera
Water boatmen have also been found in a place called Gualo.
Other insects
Plants
A lot of plant fossils have been discovered here, including tiny plant pieces and larger plant parts. These plants belong to many different groups, such as Calamitales, Coniferales, Cycadales, Equisetales, Osmundales, and Rhexoxilales.
Seed ferns
Other gymnosperms
Other plants
| Actinopterygians (ray-finned fish) of the Los Rastros Formation | |||||
|---|---|---|---|---|---|
| Genus | Species | Locality | Material | ||
| Challaia | C. elongata | Agua de la Peña Creek (San Juan Province) | A very large (>70 cm) specimen which is mostly complete, missing the tail and the front of the skull. | ||
| Gualolepis | G. carinaesquamosa | Cañón del Gualo (La Rioja Province) | Two specimens, each representing portions of the skull and anterior torso. | ||
| Rastrolepis | R. latipinnata | Cañón del Gualo (La Rioja Province) | A large (>39 cm) specimen which is mostly complete, missing the rear of the tail and the front of the skull. | ||
| R. riojaensis | Cañón del Gualo (La Rioja Province) | A large (>54 cm) specimen which is mostly complete. | |||
| Blattoptera (cockroaches and relatives) of the Los Rastros Formation | |||||
|---|---|---|---|---|---|
| Genus | Species | Locality | |||
| Argentinoblatta | A. herbsti | Río Gualo (La Rioja Province) | |||
| Argentinoblattula | A. revelata | Río Gualo (La Rioja Province) | |||
| Condorblatta | C. lutzae | Río Gualo, Los Chañares (La Rioja Province) | |||
| Hermosablatta | H. crassatella | Río Gualo, Picos Gemelos (La Rioja Province) | |||
| H. pectinata | Río Gualo (La Rioja Province) | ||||
| Lariojablatta | L. chanarensis | Los Chañares (La Rioja Province) | |||
| Mancusoblatta | M. pulchella | Río Gualo (La Rioja Province) | |||
| Pulchellablatta | P. nana | Río Gualo (La Rioja Province) | |||
| Samaroblatta | S. corrientesina | Río Gualo (La Rioja Province) | |||
| S. gualoensis | Río Gualo (La Rioja Province) | ||||
| Triassoblatta | T. argentina | Picos Gemelos (La Rioja Province) | |||
| Coleoptera (beetles and relatives) of the Los Rastros Formation | |||||
|---|---|---|---|---|---|
| Genus | Species | Locality | |||
| Ademosyne | A. arcucciae | ||||
| A. elongatus | |||||
| A. hexacostata | |||||
| A. punctuada | |||||
| Argentinocupes | A. abdalai | ||||
| A. pulcher | Río Gualo, Los Chañares (La Rioja Province) | ||||
| Argentinosyne | A. bonapartei | Los Chañares (La Rioja Province) | |||
| A. frenguellii | |||||
| A. gonaldiae | |||||
| A. gualoensis | |||||
| A. losrastroensis | |||||
| A. rugosa | |||||
| Cardiosyne | C. elegans | ||||
| C. obesa | |||||
| Gemelina | G. triangularis | ||||
| Hemipterans (true bugs) of the Los Rastros Formation | |||||
|---|---|---|---|---|---|
| Genus | Species | Locality | Material | ||
| Apheloscyta | A. mayae | Gualo (La Rioja Province) | |||
| Dysmorphoptiloides | D. losrastroensis | Gualo (La Rioja Province) | |||
| Fulgobole | F. arcucciae | Gualo (La Rioja Province) | |||
| Gallegomorphoptila | G. acostai | Gualo (La Rioja Province) | |||
| G. breviptera | Gualo (La Rioja Province) | ||||
| G. gigantea | Gualo (La Rioja Province) | ||||
| G. pulcherrima | Gualo (La Rioja Province) | ||||
| Gualoscarta | G. obscura | Gualo (La Rioja Province) | |||
| Mesoscytina | M. forsterae | Gualo (La Rioja Province) | |||
| Popovigocimex | P. yurii | Gualo (La Rioja Province) | Forewing | ||
| Saaloscytina | S. carmonae | Los Chañares, Gualo (La Rioja Province) | |||
| Yurigocimex | Y. popovi | Gualo (La Rioja Province) | Forewing | ||
| Other insects of the Los Rastros Formation | |||||
|---|---|---|---|---|---|
| Genus | Species | Locality | Material | ||
| Argentinoglosselytrina | A. pulchella | Gualo (La Rioja Province) | |||
| Argentinoperlidium | A. rogersi | Gualo (La Rioja Province) | Forewing fragment | ||
| Frenguelliphlebia | F. labandeirai | Gualo (La Rioja Province) | Forewing fragment | ||
| Miomina | M. riojana | Los Chañares (La Rioja Province) | |||
| Notopamphagopsis | N. spp. | Gualo (La Rioja Province) | Wing fragments | ||
| "Seed fern" macrofossils of the Los Rastros Formation | |||||
|---|---|---|---|---|---|
| Genus | Species | Locality | Material | ||
| Dejerseya | D. lobata | Ischigualasto Provincial Park (San Juan Province) | Frond fragments | ||
| D. lunensis | Ischigualasto Provincial Park (San Juan Province) | Frond fragments | |||
| Dicroidium | D. argenteum | Ischigualasto Provincial Park (San Juan Province) | Frond fragments | ||
| D. crassum | Ischigualasto Provincial Park (San Juan Province) | Frond fragments | |||
| D. dubium | Ischigualasto Provincial Park (San Juan Province), Ischichuca Creek | Frond fragments | |||
| D. lancifolium | Ischigualasto Provincial Park (San Juan Province), Ischichuca Creek | Frond fragments | |||
| D. lineatum | Ischigualasto Provincial Park (San Juan Province) | Frond fragments | |||
| D. nondichotoma | Ischigualasto Provincial Park (San Juan Province) | Frond fragments | |||
| D. obtusifolium | Ischigualasto Provincial Park (San Juan Province) | Frond fragments | |||
| D. odontopteroides | Ischigualasto Provincial Park (San Juan Province), Gualo (La Rioja Province), Ischichuca Creek | Frond fragments | |||
| D. pinnis-distantibus | Ischichuca Creek | ||||
| Johnstonia | J. coriacea | Agua de la Peña Creek, Ischichuca Creek | Frond fragments | ||
| J. dutoitii | Ischigualasto Provincial Park (San Juan Province) | Frond fragments | |||
| J. stelzneriana | Ischigualasto Provincial Park (San Juan Province), Gualo (La Rioja Province) Ischichuca Creek | Frond fragments | |||
| Matatiella | M. roseta | Seed cones | |||
| Pachydermophyllum | P. papillosum | Agua de la Peña Creek. | Frond fragments | ||
| P. praecordillerae | Ischigualasto Provincial Park (San Juan Province) | Frond fragments | |||
| Peltaspermum | P. monodiscum | Agua de la Peña Creek. | Seed cones | ||
| Rhexoxylon | R. sp. | Ischichuca Creek | Fossil wood | ||
| Rochipteris | R. alexandriana | Ischigualasto Provincial Park (San Juan Province) | Frond fragments | ||
| Umkomasia | U. macleanii | Strobili (seed cones) | |||
| Xylopteris | X. argentina | Ischigualasto Provincial Park (San Juan Province), Gualo (La Rioja Province), Ischichuca Creek | Frond fragments | ||
| X. densifolia | Ischigualasto Provincial Park (San Juan Province) | Frond fragments | |||
| X. elongata | Ischigualasto Provincial Park (San Juan Province), Gualo (La Rioja Province), Ischichuca Creek | Frond fragments | |||
| X. rigida | Ischigualasto Provincial Park (San Juan Province) | Frond fragments | |||
| Zuberia | Z. brownii | Ischichuca Creek | |||
| Z. feistmantelii | Gualo (La Rioja Province) | Frond fragments | |||
| Z. sahnii | Ischigualasto Provincial Park (San Juan Province), Ischichuca Creek | Frond fragments | |||
| Z. zuberi | Ischigualasto Provincial Park (San Juan Province), Ischichuca Creek | Frond fragments | |||
| Gymnosperm macrofossils of the Los Rastros Formation | |||||
|---|---|---|---|---|---|
| Genus | Species | Locality | Material | ||
| Acevedoa | A. rastroensis | Ischigualasto Provincial Park (San Juan Province) | Female fructifications | ||
| Baiera | B. africana | Ischigualasto Provincial Park (San Juan Province) | Leaf fragments | ||
| B. pontifolia | Ischigualasto Provincial Park (San Juan Province), Gualo (La Rioja Province) | Leaf fragments | |||
| B. schenkii | Ischigualasto Provincial Park (San Juan Province) | Leaf fragments | |||
| Cordaicarpus | C. sp. | Gualo (La Rioja Province) | Seeds | ||
| Cycadocarpium | C. andium | Agua de la Peña Creek. | |||
| Czekanowskia | C. rigali | Agua de la Peña Creek. | |||
| Desmiophyllum | D. sp. | Agua de la Peña Creek. | |||
| Dordrectites | D. elongatus | Ischichuca Creek | |||
| Harringtonia | H. argentinica | Agua de la Peña Creek. | |||
| Heidiphyllum | H. clarifolium | ||||
| H. elongatum | Ischigualasto Provincial Park (San Juan Province), Gualo (La Rioja Province) | Leaf fragments | |||
| H. minutifolium | |||||
| Kurtziana | K. cachuetensis | Ischigualasto Provincial Park (San Juan Province) | Frond fragments | ||
| Pelourdea | P. polyphylla | Agua de la Peña Creek. | |||
| P. problematica | Agua de la Peña Creek. | ||||
| Phoenicopsis | P. sp. | Ischichuca Creek | |||
| Pterorrachis | P. ambigua | Agua de la Peña Creek. | |||
| Rissikia | R. media | Ischigualasto Provincial Park (San Juan Province), Gualo (La Rioja Province) | Leaf fragments | ||
| Samaropsis | S. sp. | Gualo (La Rioja Province) | Seeds | ||
| Sphenobaiera | S. argentinae | Ischigualasto Provincial Park (San Juan Province) | Leaf fragments | ||
| S. sp. cf. S. argentinae | Gualo (La Rioja Province) | Leaf fragments | |||
| S. insecta | Ischigualasto Provincial Park (San Juan Province) | Leaf fragments | |||
| S. sectina | Ischigualasto Provincial Park (San Juan Province) | Leaf fragments | |||
| Sphenopteris | S. sp. | Ischichuca Creek | |||
| Telemachus | T. elongatus | Ischichuca Creek | Seed cones. | ||
| Yabeiella | Y. brackebuschiana | Ischigualasto Provincial Park (San Juan Province) | Leaf fragments | ||
| Y. mareyesiaca | |||||
| Y. spathulata | |||||
| Non-gymnosperm plant macrofossils of the Los Rastros Formation | |||||
|---|---|---|---|---|---|
| Genus | Species | Locality | Material | ||
| Cladophlebis | C. kurtzi | Agua de la Peña Creek. | |||
| C. mendozaensis | Ischigualasto Provincial Park (San Juan Province), Agua de la Peña Creek. | Frond fragments | |||
| C. mesozoica | Agua de la Peña Creek. | ||||
| C. sp. cf. C. mesozoica | Gualo (La Rioja Province) | Frond fragments | |||
| Equisetites | E. fertilis | Agua de la Peña Creek. | |||
| Neocalamites | N. carrerei | Ischichuca Creek. | |||
| N. ischigualasti | Agua de la Peña Creek. | ||||
| N. ramaccionii | Agua de la Peña Creek. | ||||
| N. sp. | Ischigualasto Provincial Park (San Juan Province), Gualo (La Rioja Province), Ischichuca Creek. | Stalk impressions | |||
| Nododendron | N. sp. | Ischichuca Creek. | |||
| Phyllotheca | P. australis | Agua de la Peña Creek. | |||
| Pleuromeia | P. sp. | Ischichuca Creek. | |||
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